Recent In-House Publications Citing Specimens from the SDSU Biodiversity Museum

Authors’ Names are Indicated as:

SDSU Faculty: bold
PostDoc: underlined
Grad Student: *
Undergrad Student: **


  • Bougie, T*., A. Brelsford, and M. Hedin. 2024. High sexual display trait diversity without measured genetic divergence in a montane hybrid zone involving young species (Habronattus americanus subgroup, Araneae: Salticidae). Insect Systematics and Diversity 8 (1) 3.


  • Azevedo, G. H. F., J. Blair and M. Hedin. 2023. Evaluating possible anthropogenic impacts on gene flow and loss of genetic diversity in endangered Madla Cave Meshweaver spiders (Hahniidae, Cicurina madla). Conservation Genetics (2023).
  • Demery, A-J*. and K. J. Burns. 2023. Widespread convergent morphological evolution within the largest family of songbirds. Evolution 77 (3) 812-822.
  • Hedin, M, and M. A. Milne. 2023. New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains. ZooKeys 1145: 1-130
  • Monjaraz-Ruedas, R., R. W. Mendez, and M. Hedin. 2023 Species delimitation, biogeography, and natural history of dwarf funnel web spiders (Mygalomorphae, Hexurellidae, Hexurella) from the United States / Mexico borderlands. ZooKeys 1167: 109-157.
  • Myers, B. M.*, K. J. Burns, C. J. Clark, and A. Brelsford. 2023. Sampling affects population genetic inference: A case study of the Allen’s (Selasphorus sasis) and Rufous Hummingbird (Selasphorus rufus). Journal of Heredity 114(6):625-636.
  • Scott, B. F*, A. J. Shultz, and K. J. Burns. 2023. The impact of habitat and migration on plumage colour in Cardinalidae. Biological Journal of the Linnean Society, blad066.
  • Simpson, M. G. and L. Gross. 2023. Noteworthy Collection: Cryptantha crassisepala (Torrey & A.Gray) Greene var. elachantha I.M.Johnston. Madroño 70: 4–8.


  • Azevedo, G. H. F., T. Bougie*, M. Carboni, M. Hedin, and M. J. Ramírez.2022. Convergence, hemiplasy and correlated evolution shape morphological diversity related to web-less lifestyle in the two-clawed spiders. Insect Systematics & Diversity 6(5): 1-14
  • Azevedo, G. H. F., T. Bougie*, M. Carboni, M. Hedin, and M. J. Ramírez.2022. Combining genomic, phenotypic and Sanger sequencing data to elucidate the phylogeny of the two-clawed spiders (Dionycha). Molecular Phylogenetics and Evolution 166: 107327
  • Ciaccio, E.*, A. Debray**, and M. Hedin. 2022. Phylogenomics of paleoendemic lampshade spiders (Araneae, Hypochilidae, Hypochilus), with the description of a new species from montane California. ZooKeys 1086: 163-204
  • Derkarabetian, S.*J. Starrettand M. Hedin. 2022. Using natural history to guide supervised machine learning for cryptic species delimitation with genetic data. Frontiers in Zoology 19:8
  • Derkarabetian, S., P. Paquin, J. Reddell, and M. Hedin. 2022. Conservation genomics of federally endangered Texella harvester species (Arachnida, Opiliones, Phalangodidae) from cave and karst habitats of central Texas. Conservation Genetics (2022)
  • Simpson, M. G., R. Vanderhoff, J. Bailey, F. Roberts, M. Mulligan, J. P. Rebman, N. Walsh, and M. Koekemoer. 2023. Taxonomic identity of a recently naturalized Senecio species in California. Madroño 69: 286–298.
  • Moroni, P. and M. G. Simpson. 2022. Revisión taxonómica del genero Johnstonella (Boraginaceae s. str.) en la Argentina. Darwiniana Nueva Serie 10: 260–270.
  • Rebman, J. P. and M. G. Simpson. 2022. Three new species of Cryptantha (Boraginaceae) from the Southern Channel Islands of California. Madroño 69: 111-134.
  • Simpson, M. G., K. Hasenstab-Lehman, M. Mabry, and M. Muñoz. 2022. Johnstonella punensis (Boraginaceae), a new species endemic to the dry Puna of Chile. PhytoKeys 197: 149–164.


  • Bougie, T. C.*, A. Brelsford, and M. Hedin. 2021. Evolutionary impacts of introgressive hybridization in a rapidly evolving group of jumping spiders (F. Salticidae, Habronattus americanus group), Molecular Phylogenetics and Evolution 161: 107165.
  • Demery, A.-J. C.*, K. J. Burns, and N. A. Mason. 2021. Bill size, bill shape, and body size constrain bird song evolution on a macroevolutionary scale. Ornithology 138 (2) ukab11.
  • Derkarabetian, S., C. M. Baker, M. Hedin, C. E. Prieto and G. Giribet. 2021. Phylogenomic reevaluation of Triaenonychoidea (Opiliones : Laniatores), and systematics of Triaenonychidae, including new families, genera, and species. Invertebrate Systematics 35: 133–157
  • Ledford, J., S. Derkarabetian, C. Griswold, C. Ribera and M. Hedin. 2021. Phylogenomics and Holarctic biogeography of leptonetid spiders (Araneae, Leptonetidae). Invertebrate Systematics 35 (3): 332-349.
  • Myers, B. M.*, K. J. Burns, C. J. Clark, and A. Brelsford. 2021. The population genetics of non-migratory Allen’s Hummingbird (Selasphorus sasin sedentarius) following a recent mainland colonization. Ecology and Evolution 11 (4): 1850-1865.
  • Ramírez, M. J., I. L. F. Magalhaes, S. Derkarabetian*, J. Ledford, C. E. Griswold, H. M. Wood and M. Hedin. 2021. Sequence-capture phylogenomics of true spiders reveals convergent evolution of respiratory systems. Systematic Biology 70 (1): 14–20.
  • Rivera, C., M. Hedin, A. C. Mason, W. P. Maddison and D. O. Elias. 2021 Complex courtship in the Habronattus clypeatus group (Araneae, Salticidae). The Journal of Arachnology 48 (3): 221-232.
  • Simpson, M. G. and J. P. Rebman. 2021. A new species of Cryptantha restricted to dunes in northwestern Baja California, Mexico. Madroño 68: 127-137.
  • Simpson, M. G. and J. P. Rebman. 2021. Research in Boraginaceae: A new variety of Cryptantha maritima, Cryptantha pondii resurrected, and Johnstonella echinosepala transferred back to Cryptantha. Phytotaxa 509: 185–210.
  • Yost, J. M., K. D. Pearson, J. Alexander, E. Gilbert, L. A. Hains, T. Barry, R. Bencie, P. Bowler, B. Carter, R. E. Crowe, E. Dean, J. Der, A. Fisher, K. Fisher, L. Flores-Renteria, C. M. Guilliams, C. Hatfield, L. Hendrickson, T. Huggins, L. Janeway, C. Lay, A. Litt, S. Markos, S. J. Mazer, D. Mccamish, L. Mcdade, M. Mesler, B. Mishler, M. Nazaire, J. Rebman, L. Rosengreen, P. W. Rundel, D. Potter, A. Sanders, K. C. Seltmann, M. G. Simpson, G. A. Wahlert, K. Waselkov, K. Williams, and P. S. Wilson. 2020. The California Phenology Collections Network: Using digital images to investigate phenological change in a biodiversity hotspot. Madroño 66: 130-141.


  • Hedin, M., S. Foldi* and B. Rajah-Boyer*. 2020. Evolutionary divergences mirror Pleistocene paleodrainages in a rapidly-evolving complex of oasis-dwelling jumping spiders (Salticidae, Habronattus tarsalis). Molecular Phylogenetics and Evolution 144: 106696.
  • Opatova, V., C. A. Hamilton, M. Hedin, L. Montes de Oca, J. Král, C. R. Haddad and J. E. Bond. 2020. Systematics and classification of the spider infraorder Mygalomorphae using genomic scale data. Systematic Biology 69: 671–707.
  • Price-Waldman, R. M.*, Shultz, A. J., and K. J. Burns. 2020. Speciation rates are correlated with changes in plumage color complexity in the largest family of songbirds. Evolution. 74 (6), 1155-1169.
  • Simpson, M. G. and R. B. Kelley. 2020. Cryptantha. Pp. 394-403 in T. J. Stephen C. Meyers, Katie E. Mitchell, Tanya Harvey   Linda K. Hardison (ed.), Flora of Oregon. Volume 2: Dicots A-F. Brit Press, Botanical Research Institute of Texas, Fort Worth, Texas.
  • Simpson, M. G., J. Stephens, and S. Yang. 2020. Noteworthy Collections: Johnstonella angelica (I.M.Johnston) Hasenstab & M.G.Simpson. Madroño 67: 5-8.


  • Benet-Pierce, N. and M. G. Simpson. 2019. The taxonomy of Chenopodium hians, C. incognitum, and ten new taxa within the narrow-leaved Chenopodium group in western North America, with special attention to California. Madroño 66: 56–75.
  • Campillo, L. C., K. J. Burns, R. G. Moyle, and J. D. Manthey. 2019. Mitochondrial genomes of the bird genus Piranga: rates of sequence evolution, and discordance between mitochondrial and nuclear markers. Mitochondrial DNA Part B 4: 2566-2569.
  • Derkarabetian, S.*, S. Castillo**, P. K. Koo, S. Ovchinnikov and M. Hedin. 2019. A demonstration of unsupervised machine learning in species delimitation. Molecular Phylogenetics and Evolution 139: 106562.
  • Hedin, M., S. Derkarabetian*, A. Alfaro**, M. J. Ramírez and J.E. Bond. 2019. Phylogenomic analysis and revised classification of atypoid mygalomorph spiders (Araneae, Mygalomorphae), with notes on arachnid ultraconserved element loci. PeerJ 7: e6864.
  • Myers, B. M.*, D. T. Rankin, K. J. Burns, and C. J. Clark. 2019. Behavioral and morphological evidence of an Allen’s × Rufous Hummingbird (Selasphorus sasin × S. rufus) hybrid zone in southern Oregon and northern California. The Auk: Ornithological Advances 136 (4) ukz049.
  • Simpson, M. G., M. E. Mabry, and K. Hasenstab-Lehman. 2019. Transfer of four species of Cryptantha to the genus Johnstonella (Boraginaceae). Phytotaxa 425: 279–289.